ESPN NFL

ENGLEWOOD, Colo. -- The Denver Broncos' first loss under rookie coach Josh McDaniels produced plenty of problems that need fixing and one big injury they'll have to deal with.

McDaniels said right tackle Ryan Harris won't play against Pittsburgh next week after sustaining a toe injury in the Broncos' loss at Baltimore. McDaniels was still trying to learn the extent of Harris' injury.

Harris, a third-year pro from Notre Dame, has started 22 consecutive games for one of the league's best offensive lines. He was replaced in the first half Sunday by second-year pro Tyler Polumbus, who is in line to make his first NFL start against the Steelers next Monday night.

Polumbus, an undrafted second-year pro from the University of Colorado, also spelled Harris in Week 3 when he went out with a shoulder injury in the second half against Oakland.

"Tyler went in and did a decent job, hung in there. Again, they've got some really good pass-rushers off the edge. He was matched up on [Ravens linebacker Terrell] Suggs on more than a few plays and handled the bull rush and speed rushers and those kinds of things," McDaniels said.

"But Tyler's been in this system as long as anybody else has this year, so he's familiar with what we're doing. We always give him plenty of reps in practice. He handled himself well in the preseason. We'll expect him to go in there and play well."

Copyright 2009 by The Associated Press